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From #3177 http://redmine.gc.cuny.edu/issues/3177#note-3

One other thing to note is bbPress has the concept of dynamic user roles. At the moment, this conflicts with Reply By Email because I wanted to copy bbPress' validation schema. And since it is hard to determine these dynamic roles when the user isn't logged in, replies will fail to post. (Or, I'm just not looking in the right place ;)

A way around this is to set the user role automatically in bbPress 2 so a role is saved in the database.

Ray - setting user role automatically is fine with me, as long as it doesn't cause other problems. I mention this because of https://bbpress.trac.wordpress.org/ticket/2210, which is the one area of bbPress that I am familiar with :)

So, it could be that this ticket can be resolved with a simple settings toggle. In any case, it's a placeholder to remind us to check.